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Pro Face-Off is a type of multiplayer mode, where both players play the same song, on the same difficulty, using the exact same notes. When playing Pro Face-Off, a meter sits in the center of the screen, between the two note highways. Initially, a needle sits at the center of the meter, and as the song progresses, the needle leans towards the player with the current highest score.
A video of a Pro Face-Off: http://www.youtube.com/watch?v=JzX4Id4eO5E